diff --git a/.travis.yml b/.travis.yml new file mode 100644 index 0000000..6aeb6ca --- /dev/null +++ b/.travis.yml @@ -0,0 +1,36 @@ +language: perl + +sudo: false + +addons: + apt: + packages: + - hmmer + - bedtools + +perl: + - "5.26" + +install: + - "export PATH=$PWD/bin:$PATH" + - "sed -i~ -e 's/-name+/-name/' bin/barrnap" + +script: + - "barrnap --version" + - "barrnap --help" + - "barrnap --citation" + - "! barrnap --doesnotexist" + - "barrnap 2>&1 | grep 'ERROR: No input file'" + - "barrnap -q --kingdom bac examples/bacteria.fna" + - "barrnap -q --kingdom arc examples/bacteria.fna" + - "barrnap -q --kingdom mito examples/mitochondria.fna" + - "barrnap -q --kingdom euk examples/fungus.fna" + - "! barrnap examples/empty.fna" + - "! barrnap examples/null.fna" + - "barrnap -q examples/small.fna | grep 16S_rRNA" + - "barrnap -q < examples/small.fna | grep 16S_rRNA" + - "barrnap -q - < examples/small.fna | grep 16S_rRNA" + - "barrnap examples/nohits.fna 2>&1 | grep 'Found 0 '" + - "barrnap --threads 2 examples/small.fna" + - "barrnap -q --incseq examples/small.fna | grep '^>'" + - "barrnap -q --outseq hits.fa < examples/small.fna && head -n3 hits.fa" diff --git a/README.md b/README.md index 8ad8587..02af920 100644 --- a/README.md +++ b/README.md @@ -1,73 +1,113 @@ +[![Build Status](https://travis-ci.org/tseemann/barrnap.svg?branch=master)](https://travis-ci.org/tseemann/barrnap) [![License: GPL v3](https://img.shields.io/badge/License-GPL%20v3-blue.svg)](https://www.gnu.org/licenses/gpl-3.0) [](#lang-au) + # Barrnap BAsic Rapid Ribosomal RNA Predictor - -## Author - -Torsten Seemann - torsten.seemann@gmail.com - @torstenseemann ## Description Barrnap predicts the location of ribosomal RNA genes in genomes. It supports bacteria (5S,23S,16S), archaea (5S,5.8S,23S,16S), -mitochondria (12S,16S) and eukaryotes (5S,5.8S,28S,18S). +metazoan mitochondria (12S,16S) and eukaryotes (5S,5.8S,28S,18S). It takes FASTA DNA sequence as input, and write GFF3 as output. -It uses the new NHMMER tool that comes with HMMER 3.1 for HMM searching in RNA:DNA style. -NHMMER binaries for 64-bit Linux and Mac OS X are included and will be auto-detected. +It uses the new `nhmmer` tool that comes with HMMER 3.1 for HMM searching in RNA:DNA style. Multithreading is supported and one can expect roughly linear speed-ups with more CPUs. -## Download +## Installation -* Tarballs: https://github.com/tseemann/barrnap/releases/ -* Source: https://github.com/tseemann/barrnap +### Requirements +* [Perl 5.xx](https://dev.perl.org/perl5/) (core modules only) +* [nhmmer](https://hmmer.org/) (part of HMMER 3.x) +* [bedtools](http://bedtools.readthedocs.io/en/latest/) -## Install +### Conda +Install [Conda](https://conda.io/docs/) or [Miniconda](https://conda.io/miniconda.html): +``` +conda -c bioconda -c conda-forge install barrnap +barrnap --version +``` - % cd $HOME - % tar zxvf barrnap-0.X.tar.gz - % echo "PATH=$PATH:$HOME/barrnap-0.x/bin" >> .bashrc - (logout and log back in) +### Homebrew +Install [HomeBrew](http://brew.sh/) (Mac OS X) or [LinuxBrew](http://brew.sh/linuxbrew/) (Linux). +``` +brew install brewsci/bio/barrnap +barrnap --help +``` + +### Source +This will install the latest version direct from Github. +You'll need to add the `bin` directory to your PATH. +``` +cd $HOME +tar zxvf barrnap-0.X.tar.gz +barrnap-0.X/barrnap -h +``` ## Usage - % barrnap --quiet examples/small.fna - ##gff-version 3 - P.marinus barrnap:0.7 rRNA 353314 354793 0 + . Name=16S_rRNA;product=16S ribosomal RNA - P.marinus barrnap:0.7 rRNA 355464 358334 0 + . Name=23S_rRNA;product=23S ribosomal RNA - P.marinus barrnap:0.7 rRNA 358433 358536 7.5e-07 + . Name=5S_rRNA;product=5S ribosomal RNA +``` +% barrnap --quiet examples/small.fna +##gff-version 3 +P.marinus barrnap:0.8 rRNA 353314 354793 0 + . Name=16S_rRNA;product=16S ribosomal RNA +P.marinus barrnap:0.8 rRNA 355464 358334 0 + . Name=23S_rRNA;product=23S ribosomal RNA +P.marinus barrnap:0.8 rRNA 358433 358536 7.5e-07 + . Name=5S_rRNA;product=5S ribosomal RNA - % barrnap -q -k mito examples/mitochondria.fna - ##gff-version 3 - AF346967.1 barrnap:0.7 rRNA 643 1610 . + . Name=12S_rRNA;product=12S ribosomal RNA - AF346967.1 barrnap:0.7 rRNA 1672 3228 . + . Name=16S_rRNA;product=16S ribosomal RNA +% barrnap -q -k mito examples/mitochondria.fna +##gff-version 3 +AF346967.1 barrnap:0.8 rRNA 643 1610 . + . Name=12S_rRNA;product=12S ribosomal RNA +AF346967.1 barrnap:0.8 rRNA 1672 3228 . + . Name=16S_rRNA;product=16S ribosomal RNA + +% barrnap -o rrna.fa < contigs.fa > rrna.gff +% head -n 3 rrna.fa +>16S_rRNA::gi|329138943|tpg|BK006945.2|:455935-456864(-) +ACGGTCGGGGGCATCAGTATTCAATTGTCAGAGGTGAAATTCTTGGATT +TATTGAAGACTAACTACTGCGAAAGCATTTGCCAAGGACGTTTTCATTA +``` + +## Options + +### General +* `--help` show help and exit +* `--version` print version in form `barrnap X.Y` and exit +* `--citation` print a citation and exit +### Search +* `--kingdom` is the database to use: Bacteria:`bac`, Archaea:`arc`, Eukaryota:`euk`, Metazoan Mitochondria:`mito` +* `--threads` is how many CPUs to assign to `nhmmer` search +* `--evalue` is the cut-off for `nhmmer` reporting, before further scrutiny +* `--lencutoff` is the proportion of the full length that qualifies as `partial` match +* `--reject` will not include hits below this proportion of the expected length +### Output +* `--quiet` will not print any messages to `stderr` +* `--incseq` will include the full input sequences in the output GFF +* `--outseq` creates a FASTA file with the hit sequences ## Caveats Barrnap does not do anything fancy. It has HMM models for each different rRNA gene. They are built from full length seed alignments. -## Requirements - -* Perl >= 5.6 -* HMMER >= 3.1b - -## License - -Barrnap is free software, released under the GPL (version 3). - ## Comparison with RNAmmer -Barrnap is designed to be a substitute for RNAmmer. It was motivated by -my desire to remove Prokka's dependency on RNAmmer -which is encumbered by a free-for-academic sign-up license, and by RNAmmer's -dependence on legacy HMMER 2.x which conflicts with HMMER 3.x that most people are using now. +Barrnap is designed to be a substitute for [RNAmmer](http://www.cbs.dtu.dk/services/RNAmmer/). +It was motivated by my desire to remove [Prokka's](https://github.com/tseemann/prokka) +dependency on RNAmmer which is encumbered by a free-for-academic sign-up +license, and by RNAmmer's dependence on legacy HMMER 2.x which conflicts +with HMMER 3.x that most people are using now. -RNAmmer is more sophisticated than Barrnap, and more accurate because it uses HMMER 2.x in glocal alignment mode whereas NHMMER 3.x currently only supports local alignment (Sean Eddy expects glocal to be supported in 2014). In practice, Barrnap will find all the typical rRNA genes in a few seconds (in bacteria), but may get the end points out by a few bases and will probably miss wierd rRNAs. The HMM models it uses are derived from Rfam, Silva and RefSeq. +RNAmmer is more sophisticated than Barrnap, and more accurate because it +uses HMMER 2.x in glocal alignment mode whereas NHMMER 3.x currently only +supports local alignment (Sean Eddy expected glocal to be supported in 2014, +but it still isn't available in 2018). + +In practice, Barrnap will find all the typical rRNA genes in a few seconds +(in bacteria), but may get the end points out by a few bases and will +probably miss wierd rRNAs. The HMM models it uses are derived from Rfam, +Silva and RefSeq. ## Data sources for HMM models -
+```
 Bacteria (70S)  
         LSU 50S
                 5S      RF00001
@@ -90,12 +130,16 @@
                 28S     SILVA-LSU-Euk
         SSU 40S
                 18S     RF01960
-        Mito
+
+Metazoan Mito
                 12S     RefSeq (MT-RNR1, s-rRNA, rns)
                 16S     RefSeq (MT-RNR2, l-rRNA, rnl)       
+```
 
-TODO: [Sajeet Haridas]
-Fungi
+## Models I would like to add
+
+```
+Fungi	[Sajeet Haridas]
         LSU 35S ?
                 5S
                 5.8S
@@ -106,16 +150,16 @@
                 15S 
                 21S (multiple exons)
                 
-
-TODO:
 Apicoplast [http://www.ncbi.nlm.nih.gov/nuccore/U87145.2]
                 LSU ~2500bp 28S ?
                 SSU ~1500bp 16S ?
 
-Plastid [Shaun Jackman]
-	?
-
-
+Plant [Shaun Jackman] + Mito [https://www.ncbi.nlm.nih.gov/nucleotide?cmd=Retrieve&dopt=GenBank&list_uids=26556996] + 5S ~118 bp ? rrn5 (use RF00001 ?) + 18S ~1935 bp ? rrn18 (use RF01960 ?) + 26S ~2568 bp ? rrn26 +``` ## Where does the name come from? @@ -125,3 +169,13 @@ The project was originally spawned at CodeFest 2013 in Berlin, Germany by Torsten Seemann and Tim Booth. +## License + +* Barrnap: [GPLv3](https://raw.githubusercontent.com/tseemann/barrnap/master/LICENSE) +* Rfam: [CC0](https://raw.githubusercontent.com/tseemann/barrnap/master/LICENSE.Rfam) +* SILVA: [Free for academic use](https://raw.githubusercontent.com/tseemann/barrnap/master/LICENSE.SILVA) + +## Author + +Torsten Seemann + diff --git a/bin/barrnap b/bin/barrnap index ec39bcb..47f7142 100755 --- a/bin/barrnap +++ b/bin/barrnap @@ -1,17 +1,17 @@ #!/usr/bin/env perl use strict; use warnings; -use Time::Piece; use List::Util qw(max); use FindBin; +use File::Temp; # . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . # global variables +my $VERSION = "0.9"; my $EXE = $FindBin::RealScript; -my $VERSION = "0.8"; my $DESC = "rapid ribosomal RNA prediction"; -my $AUTHOR = 'Torsten Seemann '; +my $AUTHOR = 'Torsten Seemann'; my $URL = 'https://github.com/tseemann/barrnap'; my $DBDIR = "$FindBin::RealBin/../db"; my $OPSYS = $^O; @@ -27,7 +27,7 @@ # . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . # command line options -my(@Options, $quiet, $kingdom, $threads, $evalue, $lencutoff, $reject, $incseq); +my(@Options, $quiet, $kingdom, $threads, $evalue, $lencutoff, $reject, $incseq, $outseq); setOptions(); # . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . @@ -40,10 +40,8 @@ msg("Adding $BINDIR to end of PATH"); $ENV{PATH} .= ":$BINDIR"; -my($NHMMER) = qx(which -a nhmmer 2> /dev/null); -$NHMMER or err("Could not find 'nhmmer' executable in PATH"); -chomp $NHMMER; -msg("Using HMMER binary: $NHMMER"); +msg("Checking for dependencies:"); +require_exe('nhmmer', 'bedtools'); $threads > 0 or err("Invalid --threads $threads"); msg("Will use $threads threads"); @@ -58,27 +56,43 @@ msg("Will reject genes < $reject of expected length."); my $kdom = $KINGDOM{ lc substr($kingdom,0,1) } or - err("I don't recognise --kingdom '$kingdom'. Try: bac arc euk mito"); + err("I don't recognise --kingdom '$kingdom'. Try:", values(%KINGDOM) ); my $hmmdb = "$DBDIR/$kdom.hmm"; err("Can't find database: $hmmdb") unless -r $hmmdb; msg("Using database: $hmmdb"); # . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . +# check if user is piping to STDIN +# nhmmer needs to fseek() so we make a temp fasta file + +my $fasta = shift @ARGV; +my $tmpfh; +if (defined($fasta) && $fasta eq '-' or !defined($fasta) && !-t \*STDIN) { + $tmpfh = File::Temp->new(UNLINK=>1); + msg("Copying STDIN to a temporary file:", $tmpfh->filename); + while () { + print $tmpfh $_; + } + $fasta = $tmpfh->filename; +} +$fasta && -r $fasta or err("No input file on command line or stdin"); + +# . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . # run the external command -my $fasta = shift @ARGV; -$fasta && -r $fasta or err("Usage: $EXE "); - msg("Scanning $fasta for $kdom rRNA genes... please wait"); -my $cmd = "$NHMMER --cpu $threads -E $evalue --w_length $MAXLEN -o /dev/null --tblout /dev/stdout \Q$hmmdb\E \Q$fasta\E"; +my $opts = "--cpu $threads -E $evalue --w_length $MAXLEN -o /dev/null --tblout /dev/stdout"; +my $cmd = "nhmmer $opts '$hmmdb' '$fasta'"; msg("Command: $cmd"); my @hits = qx($cmd 2>&1); # . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . # process the output +my %hitname; my @feat; +my @bed; HIT: foreach (@hits) { chomp; @@ -88,9 +102,13 @@ my @x = split ' ', $_; err("bad line in nhmmer output - @x") unless defined $x[6] and $x[6] =~ m/^\d+$/; + # massage to GFF my($begin,$end,$strand) = $x[6] < $x[7] ? ($x[6],$x[7],'+') : ($x[7],$x[6],'-'); my($seqid, $gene, $prod) = ($x[0], $x[2], $x[2]); my $score = defined $x[12] ? $x[12] : '.'; + + # record hits for --outseq retrieval later + $hitname{"$seqid/$begin:$end($strand)"} = $prod; # check if hit makes sense to us exists $LENG{$gene} or err("Detected unknown gene '$gene' in scan, aborting."); @@ -112,6 +130,9 @@ $prod .= " (partial)"; } + # keep track of good hits for retrievel later + push @bed, [ $seqid, $begin-1, $end, $gene, 100, $strand ]; + msg("Found:", $gene, $seqid, "L=$len/$LENG{$gene}", "$begin..$end", $strand, $prod); my $tags = "Name=$gene;product=$prod"; $tags .= ";note=$note" if $note; @@ -140,38 +161,68 @@ print while (); # `cat $fasta` } -#---------------------------------------------------------------------- - +if ($outseq) { + msg("Writing hit sequences to: $outseq"); + my $bed = File::Temp->new(); + for my $b (@bed) { + print $bed join("\t", @$b),"\n"; + } + $bed->seek(0, SEEK_END); # rewind + my $cmd = "bedtools getfasta -s -name+ -fo '$outseq' -fi '$fasta' -bed '".$bed->filename."'"; + msg("Running: $cmd"); + system($cmd); +} + +# . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . +# final cleanup + +msg("Done."); + +#---------------------------------------------------------------------- +sub require_exe { + for my $exe (@_) { + my($which) = qx(which $exe 2> /dev/null); + $which or err("Can not find required '$exe' in PATH"); + chomp $which; + msg("Found $exe - $which"); + } +} + +#---------------------------------------------------------------------- +sub revcom { + my($s) = @_; + $s = reverse($s); + $s =~ tr/ATCGatcg/TAGCtagc/; + return $s; +} + +#---------------------------------------------------------------------- sub msg { return if $quiet; - my $t = localtime; - my $line = "[".$t->hms."] @_\n"; + my $line = "[$EXE] @_\n"; print STDERR $line; } #---------------------------------------------------------------------- - sub err { $quiet=0; - msg(@_); + msg("ERROR:", @_); exit(2); } #---------------------------------------------------------------------- - sub version { print STDERR "$EXE $VERSION\n"; exit; } #---------------------------------------------------------------------- - sub show_citation { print STDERR << "EOCITE"; If you use Barrnap in your work, please cite: - Seemann T (2013) + Seemann T $EXE $VERSION : $DESC $URL @@ -193,19 +244,19 @@ {OPT=>"help", VAR=>\&usage, DESC=>"This help"}, {OPT=>"version", VAR=>\&version, DESC=>"Print version and exit"}, {OPT=>"citation",VAR=>\&show_citation, DESC=>"Print citation for referencing $EXE"}, - {OPT=>"kingdom=s", VAR=>\$kingdom, DEFAULT=>'bac', - DESC=>"Kingdom: ".join(' ', values %KINGDOM) }, + {OPT=>"kingdom=s", VAR=>\$kingdom, DEFAULT=>'bac', DESC=>"Kingdom: ".join(' ', values %KINGDOM) }, {OPT=>"quiet!", VAR=>\$quiet, DEFAULT=>0, DESC=>"No screen output"}, - {OPT=>"threads=i", VAR=>\$threads, DEFAULT=>8, DESC=>"Number of threads/cores/CPUs to use"}, + {OPT=>"threads=i", VAR=>\$threads, DEFAULT=>1, DESC=>"Number of threads/cores/CPUs to use"}, {OPT=>"lencutoff=f",VAR=>\$lencutoff, DEFAULT=>0.8, DESC=>"Proportional length threshold to label as partial"}, - {OPT=>"reject=f",VAR=>\$reject, DEFAULT=>0.5, DESC=>"Proportional length threshold to reject prediction"}, + {OPT=>"reject=f",VAR=>\$reject, DEFAULT=>0.25, DESC=>"Proportional length threshold to reject prediction"}, {OPT=>"evalue=f",VAR=>\$evalue, DEFAULT=>1E-6, DESC=>"Similarity e-value cut-off"}, - {OPT=>"incseq!", VAR=>\$incseq, DEFAULT=>0, DESC=>"Include FASTA input sequences in GFF3 output"}, + {OPT=>"incseq!", VAR=>\$incseq, DEFAULT=>0, DESC=>"Include FASTA _input_ sequences in GFF3 output"}, + {OPT=>"outseq=s", VAR=>\$outseq, DEFAULT=>'', DESC=>"Save rRNA hit seqs to this FASTA file"}, ); - (!@ARGV) && (usage()); - - &GetOptions(map {$_->{OPT}, $_->{VAR}} grep { ref } @Options) || usage(); +# (!@ARGV) && (usage(1)); + + &GetOptions(map {$_->{OPT}, $_->{VAR}} grep { ref } @Options) || usage(1); # Now setup default values. foreach (@Options) { @@ -218,9 +269,17 @@ #---------------------------------------------------------------------- sub usage { - print STDERR "Synopsis:\n $EXE $VERSION - $DESC\n"; - print STDERR "Author:\n $AUTHOR\n"; - print STDERR "Usage:\n $EXE [options] \n"; + my($exitcode) = @_; + $exitcode = 0 if $exitcode eq 'help'; # what gets passed by getopt func ref + $exitcode ||= 0; + select STDERR if $exitcode; # write to STDERR if exitcode is error + + print "Synopsis:\n $EXE $VERSION - $DESC\n"; + print "Author:\n $AUTHOR\n"; + print "Usage:\n"; + print " $EXE [options] chr.fa\n"; + print " $EXE [options] < chr.fa\n"; + print " $EXE [options] - < chr.fa\n"; foreach (@Options) { if (ref) { my $def = defined($_->{DEFAULT}) ? " (default '$_->{DEFAULT}')" : ""; @@ -230,13 +289,13 @@ $opt =~ s/=s$/ [X]/; $opt =~ s/=i$/ [N]/; $opt =~ s/=f$/ [n.n]/; - printf STDERR " --%-15s %s%s\n", $opt, $_->{DESC}, $def; + printf " --%-15s %s%s\n", $opt, $_->{DESC}, $def; } else { - print STDERR "$_\n"; + print "$_\n"; } } - exit(1); -} - -#---------------------------------------------------------------------- + exit($exitcode); +} + +#---------------------------------------------------------------------- diff --git a/db/euk.hmm b/db/euk.hmm index c5df87e..4287ced 100644 --- a/db/euk.hmm +++ b/db/euk.hmm @@ -1,3 +1,5581 @@ +HMMER3/f [3.1b2 | February 2015] +NAME 18S_rRNA +ACC RF01960 +DESC Eukaryotic small subunit ribosomal RNA +LENG 1851 +MAXL 2860 +ALPH RNA +RF yes +MM no +CONS yes +CS yes +MAP yes +DATE Thu Apr 20 11:07:37 2017 +NSEQ 91 +EFFN 4.429474 +CKSUM 3445497107 +STATS LOCAL MSV -13.4315 0.69488 +STATS LOCAL VITERBI -14.7858 0.69488 +STATS LOCAL FORWARD -7.0200 0.69488 +HMM A C G U + m->m m->i m->d i->m i->i d->m d->d + COMPO 1.33849 1.53541 1.34060 1.34437 + 1.38629 1.38629 1.38629 1.38629 + 0.09303 4.19411 2.60708 1.46634 0.26236 0.00000 * + 1 1.15271 1.89687 2.03589 0.90729 1 u u - : + 1.38629 1.38629 1.38629 1.38629 + 0.03257 4.13365 4.13365 1.46634 0.26236 1.04592 0.43289 + 2 0.57824 1.95456 2.05834 1.77301 2 a A - : + 1.38629 1.38629 1.38629 1.38629 + 0.03184 4.15616 4.15616 1.46634 0.26236 0.45528 1.00587 + 3 2.04131 0.67716 2.31771 1.33338 3 c C - : + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 4 3.00883 0.23904 2.96558 2.19164 4 c C - < + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 5 2.70790 2.22080 2.95522 0.25782 5 u U - < + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 6 2.45358 3.07488 0.20468 2.93911 6 g G - < + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 7 2.20330 3.13796 0.22805 2.99359 7 g G - < + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 8 3.05295 2.54038 3.20227 0.18239 8 u U - < + 1.38629 1.38629 1.38629 1.38629 + 0.03709 4.19411 3.84769 1.46634 0.26236 1.09861 0.40547 + 9 2.52990 2.13027 2.52319 0.32667 9 u U - _ + 1.38629 1.38629 1.38629 1.38629 + 0.03083 4.18785 4.18785 1.46634 0.26236 1.22789 0.34660 + 10 2.52519 3.50347 0.15915 3.29704 10 g G - _ + 1.38629 1.38629 1.38629 1.38629 + 0.03083 4.18785 4.18785 1.46634 0.26236 1.22789 0.34660 + 11 0.14048 3.20321 3.04324 3.15244 11 a A - _ + 1.38629 1.38629 1.38629 1.38629 + 0.03083 4.18785 4.18785 1.46634 0.26236 1.22789 0.34660 + 12 2.65946 2.41462 2.98321 0.23574 12 u U - _ + 1.38629 1.38629 1.38629 1.38629 + 0.03473 4.18785 3.96556 1.46634 0.26236 0.88146 0.53474 + 13 3.03671 0.25827 3.21320 1.97045 13 c C - _ + 1.38629 1.38629 1.38629 1.38629 + 0.03075 4.19035 4.19035 1.46634 0.26236 1.17833 0.36787 + 14 3.21817 0.17668 3.40515 2.42220 14 c C - _ + 1.38629 1.38629 1.38629 1.38629 + 0.03075 4.19035 4.19035 1.46634 0.26236 1.17833 0.36787 + 15 3.00919 2.46274 3.16013 0.19474 15 u U - _ + 1.38629 1.38629 1.38629 1.38629 + 0.03075 4.19035 4.19035 1.46634 0.26236 0.95604 0.48518 + 16 2.45701 3.47595 0.16813 3.26658 16 g G - > + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 17 3.07961 0.22255 3.15187 2.20026 17 c C - > + 1.41687 1.37631 1.41687 1.33732 + 0.03709 3.84769 4.19411 1.33209 0.30642 1.09861 0.40547 + 18 3.01214 0.20055 3.10157 2.43545 21 c C - > + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 19 0.24050 2.87491 2.54516 2.53977 22 a A - > + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 20 2.54497 3.09219 0.18567 3.08837 23 g G - > + 1.38629 1.38629 1.38629 1.38629 + 0.03872 4.19411 3.77694 1.46634 0.26236 1.09861 0.40547 + 21 2.10076 2.02400 2.39866 0.42362 24 u U - , + 1.38629 1.38629 1.38629 1.38629 + 0.03088 4.18627 4.18627 1.46634 0.26236 0.84063 0.56465 + 22 0.63149 1.98133 1.78400 1.81799 25 a a - { + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 23 1.78422 2.11899 0.60546 1.79519 26 g g - { + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 24 2.14750 1.75794 2.22365 0.50648 27 u u - { + 1.38629 1.38629 1.38629 1.38629 + 0.24012 4.19411 1.61754 1.46634 0.26236 1.09861 0.40547 + 25 1.98283 0.73347 1.75042 1.56839 28 c c - 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> + 1.38629 1.38629 1.38629 1.38629 + 0.03095 4.18412 4.18412 1.46634 0.26236 1.29717 0.31925 + 1028 1.81481 2.52708 0.38865 2.53497 1844 g g - > + 1.38629 1.38629 1.38629 1.38629 + 0.03095 4.18412 4.18412 1.46634 0.26236 1.29717 0.31925 + 1029 2.08007 1.43693 2.28107 0.62503 1845 u u - > + 1.38629 1.38629 1.38629 1.38629 + 0.03095 4.18412 4.18412 1.46634 0.26236 1.29717 0.31925 + 1030 2.56924 0.43485 2.38948 1.69083 1846 c C - > + 1.41952 1.34215 1.41952 1.36626 + 0.05241 3.32909 4.18412 0.70551 0.68093 1.29717 0.31925 + 1031 2.08692 1.47899 0.66061 2.02853 1848 g G - - + 1.38629 1.38629 1.38629 1.38629 + 0.03095 4.18412 4.18412 1.46634 0.26236 1.29717 0.31925 + 1032 3.20233 2.86129 3.33523 0.14326 1849 u U - - + 1.38629 1.38629 1.38629 1.38629 + 0.03095 4.18412 4.18412 1.46634 0.26236 1.29717 0.31925 + 1033 0.13059 3.46045 2.92241 3.29142 1850 a A - - + 1.38629 1.38629 1.38629 1.38629 + 0.03095 4.18412 4.18412 1.46634 0.26236 1.29717 0.31925 + 1034 2.06185 2.61225 0.36747 2.23559 1851 g g - > + 1.38629 1.38629 1.38629 1.38629 + 0.03095 4.18412 4.18412 1.46634 0.26236 1.29717 0.31925 + 1035 2.39040 2.25373 2.31837 0.34961 1852 u u - > + 1.40621 1.40621 1.40621 1.32884 + 0.06374 3.59389 3.37383 0.88546 0.53192 0.79100 0.60403 + 1036 2.05191 0.96377 1.99987 1.03646 1854 c c - > + 1.38629 1.38629 1.38629 1.38629 + 0.03123 4.17510 4.17510 1.46634 0.26236 1.08368 0.41302 + 1037 2.22079 0.72861 2.29727 1.17648 1855 c C - > + 1.32511 1.40755 1.40755 1.40755 + 0.04465 3.56156 4.18180 0.86308 0.54794 1.33791 0.30434 + 1038 1.60951 1.58375 1.98997 0.78064 1857 u u - > + 1.38629 1.38629 1.38629 1.38629 + 0.03102 4.18180 4.18180 1.46634 0.26236 1.33791 0.30434 + 1039 1.00692 1.82364 1.23121 1.70772 1858 a a - > + 1.43801 1.43801 1.43801 1.24549 + 0.06503 3.04303 4.18180 0.55072 0.85928 1.33791 0.30434 + 1040 0.48258 2.39254 1.83594 2.02532 1860 a a - > + 1.38629 1.38629 1.38629 1.38629 + 0.03102 4.18180 4.18180 1.46634 0.26236 1.33791 0.30434 + 1041 2.92839 0.26040 2.82173 2.15187 1861 c C - > + 1.38629 1.38629 1.38629 1.38629 + 0.03102 4.18180 4.18180 1.46634 0.26236 1.33791 0.30434 + 1042 2.01027 0.79016 2.23777 1.18555 1862 c c - , + 1.38629 1.38629 1.38629 1.38629 + 0.03102 4.18180 4.18180 1.46634 0.26236 1.33791 0.30434 + 1043 1.10868 1.94684 1.04506 1.73949 1863 g a - , + 1.38629 1.38629 1.38629 1.38629 + 0.03102 4.18180 4.18180 1.46634 0.26236 1.33791 0.30434 + 1044 2.87485 2.20300 3.02663 0.24255 1864 u U - , + 1.38629 1.38629 1.38629 1.38629 + 0.03102 4.18180 4.18180 1.46634 0.26236 0.74399 0.64476 + 1045 0.16806 3.30935 2.60190 3.12295 1865 a A - , + 1.32884 1.40621 1.40621 1.40621 + 0.04322 3.60388 4.19411 0.88546 0.53192 1.09861 0.40547 + 1046 0.08229 3.76409 3.53702 3.62298 1867 A A - , + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1047 0.18119 3.05986 3.00165 2.67186 1868 a A - , + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1048 2.78379 0.31011 3.06214 1.84487 1869 c C - , + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1049 1.69808 1.95262 0.97910 1.20592 1870 g g - , + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1050 0.45972 2.11059 2.29797 1.91795 1871 a A - , + 1.38629 1.38629 1.38629 1.38629 + 0.04093 4.19411 3.68793 1.46634 0.26236 1.09861 0.40547 + 1051 2.99969 2.46040 3.14993 0.19609 1872 u U - , + 1.38629 1.38629 1.38629 1.38629 + 0.03095 4.18412 4.18412 1.46634 0.26236 0.79100 0.60403 + 1052 1.55717 2.72520 0.46320 2.35948 1873 g g - ( + 1.38629 1.38629 1.38629 1.38629 + 0.03948 4.19411 3.74537 1.46634 0.26236 1.09861 0.40547 + 1053 2.14678 0.56935 2.17543 1.59117 1874 c c - ( + 1.38629 1.38629 1.38629 1.38629 + 0.03090 4.18553 4.18553 1.46634 0.26236 1.27154 0.32906 + 1054 1.88017 0.77759 1.80860 1.49592 1875 c c - ( + 1.38629 1.38629 1.38629 1.38629 + 0.04129 4.18553 3.67935 1.46634 0.26236 0.82280 0.57839 + 1055 0.99678 2.13636 1.03833 1.84007 1876 a a - ( + 1.39780 1.39780 1.39780 1.35255 + 0.03825 3.80365 4.18412 1.05832 0.42624 1.29717 0.31925 + 1056 0.71219 1.81502 1.74188 1.76372 1878 a a - ( + 1.38629 1.38629 1.38629 1.38629 + 0.03095 4.18412 4.18412 1.46634 0.26236 1.29717 0.31925 + 1057 2.36253 0.38370 2.81982 1.80261 1879 c C - ( + 1.38629 1.38629 1.38629 1.38629 + 0.03095 4.18412 4.18412 1.46634 0.26236 1.29717 0.31925 + 1058 1.70478 1.18847 2.05684 0.95284 1880 u u - , + 1.38629 1.38629 1.38629 1.38629 + 0.03095 4.18412 4.18412 1.46634 0.26236 0.79100 0.60403 + 1059 0.97525 1.78806 1.29549 1.70455 1881 a g - < + 1.38629 1.38629 1.38629 1.38629 + 0.04854 4.19411 3.43284 1.46634 0.26236 1.09861 0.40547 + 1060 1.66786 2.38214 0.54865 1.95709 1882 g g - < + 1.38629 1.38629 1.38629 1.38629 + 0.03118 4.17675 4.17675 1.46634 0.26236 0.91085 0.51447 + 1061 1.40653 1.31761 1.16005 1.75007 1883 g c - < + 1.36006 1.36006 1.43743 1.38962 + 0.05134 3.35655 4.18691 0.90848 0.51606 0.85672 0.55261 + 1062 1.45414 1.87333 0.88567 1.60769 1886 g g - - + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1063 0.89283 1.92584 1.47710 1.53038 1887 a a - - + 1.38629 1.38629 1.38629 1.38629 + 0.05587 4.19411 3.23781 1.46634 0.26236 1.09861 0.40547 + 1064 1.95353 1.86492 1.98291 0.56978 1888 u u - < + 1.38629 1.38629 1.38629 1.38629 + 0.03140 4.16965 4.16965 1.46634 0.26236 0.86221 0.54858 + 1065 1.60255 1.39105 1.74642 0.97975 1889 u u - < + 1.38629 1.38629 1.38629 1.38629 + 0.03095 4.18412 4.18412 1.46634 0.26236 1.29717 0.31925 + 1066 1.53944 1.63444 0.90526 1.68201 1890 g g - < + 1.38629 1.38629 1.38629 1.38629 + 0.03095 4.18412 4.18412 1.46634 0.26236 1.29717 0.31925 + 1067 2.28085 2.69355 0.34329 2.11421 1891 g g - < + 1.38629 1.38629 1.38629 1.38629 + 0.03095 4.18412 4.18412 1.46634 0.26236 1.29717 0.31925 + 1068 1.44467 1.39885 1.51746 1.21057 1892 u c - < + 1.38629 1.38629 1.38629 1.38629 + 0.03727 4.18412 3.84679 1.46634 0.26236 1.29717 0.31925 + 1069 1.45267 1.47204 1.32028 1.31105 1893 u c - < + 1.38629 1.38629 1.38629 1.38629 + 0.03114 4.17799 4.17799 1.46634 0.26236 1.40122 0.28276 + 1070 1.45106 1.66956 1.06199 1.46282 1894 g g - < + 1.41553 1.41553 1.36799 1.34789 + 0.05008 3.39565 4.17799 0.75116 0.63832 1.40122 0.28276 + 1071 1.31717 1.83743 1.16908 1.33854 1896 g a - < + 1.40030 1.40030 1.40030 1.34541 + 0.14448 3.73054 2.20230 0.99980 0.45879 1.40122 0.28276 + 1072 1.23857 1.62960 1.50031 1.23398 1898 u a - - + 1.38629 1.38629 1.38629 1.38629 + 0.03447 4.07793 4.07793 1.46634 0.26236 0.92372 0.50590 + 1073 1.54112 1.98432 0.80984 1.59230 1899 g g - < + 1.38629 1.38629 1.38629 1.38629 + 0.03274 4.12877 4.12877 1.46634 0.26236 1.97317 0.14968 + 1074 1.48225 1.67700 1.61141 0.95105 1900 u u - < + 1.38629 1.38629 1.38629 1.38629 + 0.03274 4.12877 4.12877 1.46634 0.26236 1.72363 0.19652 + 1075 1.58319 1.22527 1.96031 1.02113 1901 u c - < + 1.38560 1.25925 1.54639 1.37458 + 0.15790 2.03994 4.13332 0.59332 0.80406 1.44900 0.26762 + 1076 1.07562 1.51301 1.67741 1.37908 1905 a a - 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_ + 1.30986 1.76295 1.56053 1.05389 + 0.34548 1.52892 2.58565 3.15809 0.04344 0.25264 1.49947 + 1084 1.58712 1.97864 0.73863 1.71778 1975 g g - > + 1.38629 1.38629 1.38629 1.38629 + 0.03752 4.12722 3.87767 1.46634 0.26236 1.98642 0.14756 + 1085 0.86563 1.66143 1.71930 1.55992 1976 a a - > + 1.38629 1.38629 1.38629 1.38629 + 0.03294 4.12264 4.12264 1.46634 0.26236 2.02457 0.14162 + 1086 1.68220 0.96090 1.92302 1.25414 1977 c c - > + 1.38629 1.38629 1.38629 1.38629 + 0.04828 4.12264 3.47609 1.46634 0.26236 0.51767 0.90611 + 1087 1.52576 1.29634 1.76814 1.08364 1978 u u - > + 1.38629 1.38629 1.38629 1.38629 + 0.03159 4.16396 4.16396 1.46634 0.26236 0.95669 0.48477 + 1088 1.64795 1.21634 1.59917 1.17384 1979 u c - > + 1.38629 1.38629 1.38629 1.38629 + 0.04605 4.17799 3.51743 1.46634 0.26236 1.40122 0.28276 + 1089 1.48363 1.40277 1.41413 1.25830 1980 u g - > + 1.38629 1.38629 1.38629 1.38629 + 0.03160 4.16354 4.16354 1.46634 0.26236 0.75974 0.63071 + 1090 1.38227 1.54334 1.32899 1.30718 1981 u g - 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> + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1147 1.66665 2.39781 0.53842 1.99118 2079 g g - > + 1.40621 1.40621 1.32884 1.40621 + 0.04322 3.60388 4.19411 0.88546 0.53192 1.09861 0.40547 + 1148 2.16043 0.56937 2.09781 1.62904 2081 c C - > + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1149 2.04014 1.93186 2.15956 0.49473 2082 u u - > + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1150 2.41258 3.10349 0.23428 2.59855 2083 g G - - + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1151 0.34561 2.56602 2.43179 2.05971 2084 a A - - + 1.38629 1.38629 1.38629 1.38629 + 0.03701 4.19411 3.85142 1.46634 0.26236 1.09861 0.40547 + 1152 0.17792 2.76214 3.09703 2.90684 2085 a A - - + 1.38629 1.38629 1.38629 1.38629 + 0.03083 4.18793 4.18793 1.46634 0.26236 0.88365 0.53319 + 1153 0.16194 3.14435 3.16356 2.74669 2086 a A - 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, + 1.38629 1.38629 1.38629 1.38629 + 0.03134 4.17169 4.17169 1.46634 0.26236 1.49739 0.25323 + 1161 2.08398 2.97559 0.28493 2.62447 2098 g G - , + 1.40379 1.33558 1.40379 1.40379 + 0.04264 3.63762 4.17169 0.92883 0.50255 1.49739 0.25323 + 1162 0.33794 2.31290 2.48067 2.26241 2100 a A - , + 1.38629 1.38629 1.38629 1.38629 + 0.03134 4.17169 4.17169 1.46634 0.26236 1.49739 0.25323 + 1163 0.17390 3.07856 3.09788 2.68166 2101 a A - , + 1.38629 1.38629 1.38629 1.38629 + 0.03134 4.17169 4.17169 1.46634 0.26236 1.49739 0.25323 + 1164 3.17602 2.83940 3.30830 0.14710 2102 u U - , + 1.38629 1.38629 1.38629 1.38629 + 0.03134 4.17169 4.17169 1.46634 0.26236 0.59317 0.80424 + 1165 2.90475 2.65021 2.77872 0.20766 2103 u U - { + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1166 2.41826 3.45577 0.17390 3.24479 2104 g G - { + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1167 0.14337 3.41102 2.79353 3.23506 2105 a A - 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> + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1805 1.04603 2.15075 0.92536 1.99596 3225 g g - > + 1.38629 1.38629 1.38629 1.38629 + 0.04167 4.19411 3.66004 1.46634 0.26236 1.09861 0.40547 + 1806 0.16822 3.10953 3.12975 2.71086 3226 a A - - + 1.38629 1.38629 1.38629 1.38629 + 0.03097 4.18341 4.18341 1.46634 0.26236 0.77584 0.61677 + 1807 0.16362 3.13576 3.15727 2.73453 3227 a A - - + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1808 2.76330 3.63683 0.12955 3.43916 3228 g G - > + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1809 3.02439 2.48324 3.17530 0.19097 3229 u U - - + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1810 3.17614 0.19458 3.36248 2.29835 3230 c C - > + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1811 2.26740 3.12830 0.23784 2.74420 3231 g G - > + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1812 3.22346 2.87889 3.35684 0.14025 3232 u U - - + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1813 0.16362 3.13576 3.15727 2.73453 3233 a A - - + 1.38629 1.38629 1.38629 1.38629 + 0.04449 4.19411 3.56028 1.46634 0.26236 1.09861 0.40547 + 1814 0.24851 2.61946 2.84620 2.41767 3234 a A - > + 1.38629 1.38629 1.38629 1.38629 + 0.03105 4.18067 4.18067 1.46634 0.26236 1.35707 0.29761 + 1815 3.23127 0.16846 3.41652 2.49256 3235 c C - > + 1.38629 1.38629 1.38629 1.38629 + 0.03105 4.18067 4.18067 1.46634 0.26236 1.35707 0.29761 + 1816 0.08527 3.72899 3.50569 3.58668 3236 A A - - + 1.38629 1.38629 1.38629 1.38629 + 0.03105 4.18067 4.18067 1.46634 0.26236 0.72325 0.66393 + 1817 0.22560 2.59229 2.93026 2.60729 3237 a A - - + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1818 2.35597 3.18085 0.21960 2.79999 3238 g G - > + 1.36136 1.39475 1.39475 1.39475 + 0.03593 3.90183 4.19411 1.14066 0.38509 1.09861 0.40547 + 1819 3.09270 3.81113 0.09897 3.62220 3240 G G - , + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1820 2.91709 2.25773 3.07024 0.22953 3241 u U - < + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1821 1.60738 1.78698 2.24762 0.64156 3242 u u - < + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1822 1.95124 2.00133 1.81744 0.57927 3243 u u - < + 1.39258 1.36766 1.39258 1.39258 + 0.03456 3.96912 4.19411 1.20793 0.35498 1.09861 0.40547 + 1823 2.64777 0.50822 2.70749 1.34354 3245 c c - < + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1824 2.13607 0.64647 2.06640 1.46382 3246 c c - < + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1825 2.32075 3.40514 0.18943 3.19009 3247 g G - < + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1826 2.99416 2.79994 3.25628 0.16184 3248 u U - < + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1827 0.14962 3.21130 3.21140 2.84110 3249 a A - < + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1828 2.59955 3.38494 0.16831 3.06356 3250 g G - < + 1.38629 1.38629 1.38629 1.38629 + 0.05417 4.19411 3.27960 1.46634 0.26236 1.09861 0.40547 + 1829 2.74197 3.59671 0.13384 3.39884 3251 g G - < + 1.40504 1.40504 1.35581 1.38012 + 0.03538 3.94630 4.17129 1.37083 0.29289 0.58858 0.80994 + 1830 2.84045 2.65074 2.53839 0.23318 3255 u U - _ + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1831 2.72200 3.61502 0.13411 3.41594 3256 g G - _ + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1832 0.08229 3.76409 3.53702 3.62298 3257 A A - _ + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1833 0.08229 3.76409 3.53702 3.62298 3258 A A - _ + 1.38629 1.38629 1.38629 1.38629 + 0.06035 4.19411 3.13545 1.46634 0.26236 1.09861 0.40547 + 1834 3.20030 0.17358 3.38376 2.46884 3259 c C - > + 1.38629 1.38629 1.38629 1.38629 + 0.03154 4.16530 4.16530 1.46634 0.26236 0.52700 0.89251 + 1835 2.96848 0.24716 2.83991 2.21481 3260 c C - > + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1836 2.62527 2.59596 2.67772 0.24298 3261 u U - > + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1837 2.48926 2.83329 0.21555 2.95438 3262 g G - > + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1838 3.21350 0.17993 3.40088 2.39565 3263 c C - > + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1839 1.16980 2.48369 0.68079 2.30337 3264 g g - > + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1840 2.24065 3.01328 0.26880 2.52338 3265 g g - > + 1.38629 1.38629 1.38629 1.38629 + 0.03456 4.19411 3.96912 1.46634 0.26236 1.09861 0.40547 + 1841 0.45417 2.03044 2.29685 2.01612 3266 a a - > + 1.38629 1.38629 1.38629 1.38629 + 0.03075 4.19030 4.19030 1.46634 0.26236 1.17937 0.36741 + 1842 0.62086 2.11978 1.82230 1.71034 3267 a a - > + 1.38629 1.38629 1.38629 1.38629 + 0.04382 4.19030 3.58512 1.46634 0.26236 1.17937 0.36741 + 1843 2.26637 3.22608 0.21857 2.93875 3268 g G - > + 1.39717 1.39717 1.35435 1.39717 + 0.05932 3.81472 3.33679 1.07422 0.41789 0.67282 0.71390 + 1844 2.39421 2.92239 0.22071 2.93749 3270 g G - : + 1.38629 1.38629 1.38629 1.38629 + 0.03126 4.17402 4.17402 1.46634 0.26236 0.89019 0.52861 + 1845 0.23073 2.57294 2.90829 2.58805 3271 a A - : + 1.38629 1.38629 1.38629 1.38629 + 0.03089 4.18585 4.18585 1.46634 0.26236 0.83040 0.57248 + 1846 2.80908 2.34102 3.03867 0.22864 3272 u U - : + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1847 2.82705 0.20433 3.27658 2.43194 3273 c C - : + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1848 0.27554 2.55043 2.45997 2.55940 3274 a A - : + 1.38629 1.38629 1.38629 1.38629 + 0.03063 4.19411 4.19411 1.46634 0.26236 1.09861 0.40547 + 1849 1.21349 1.95745 1.89958 0.88675 3275 u u - : + 1.38629 1.38629 1.38629 1.38629 + 0.11086 4.19411 2.40957 1.46634 0.26236 1.09861 0.40547 + 1850 1.84088 1.85329 1.36425 0.84623 3276 u u - : + 1.38629 1.38629 1.38629 1.38629 + 0.05345 4.11640 3.33141 1.46634 0.26236 2.07403 0.13431 + 1851 0.93797 1.35881 1.81169 1.67006 3277 a a - : + 1.38629 1.38629 1.38629 1.38629 + 0.01705 4.08000 * 1.46634 0.26236 0.00000 * +// HMMER3/f [3.1b2 | February 2015] NAME 5S_rRNA ACC RF00001 @@ -10,7 +5588,7 @@ CONS yes CS yes MAP yes -DATE Sun Feb 19 15:32:20 2017 +DATE Thu Apr 20 11:07:36 2017 NSEQ 712 EFFN 9.307953 CKSUM 242171328 @@ -392,7 +5970,7 @@ CONS yes CS yes MAP yes -DATE Sun Feb 19 15:32:20 2017 +DATE Thu Apr 20 11:07:36 2017 NSEQ 61 EFFN 4.121521 CKSUM 863152359